Evolution (L567) 2013
Department of Biology,
evolution (L567).--This is a graduate course in
evolution, focusing on the theoretical underpinnings.
Beginning in 2010, I have put more focus on Evolutionary Ecology
and Behavioral Ecology. The goal will be to complement our
graduate courses in Evolutionary Genetics. Topics will
include life-history theory, evolution of sex, sexual selection,
evolution of social behavior, evolution of virulence, and the
evolutionary ecology of speciation and macroevolution. I also plan
to cover some of the basic theory for Population Genetics and
Quantitative Genetics. Graduate standing or permission of
the instructor is required. Advanced undergraduates in Biology are
The course will be offered
again during the Fall semester of 2015. Typically, the
class meetings are part lecture and part discussion in an
Optional (but highly recommended)
computer exercise in population genetics. Click on Pop.
Simulation Instructions to download the
worksheet. After you fill out the worksheet, you can
compare your answers to Answers to
PopGen exercise. Make sure you run through the
exercise above before looking at the answers.
for qhat (mathematica
mutation-selection balance example. Here is the solution
Population-Genetic exercise #1 due 12 Sept 2013. There
will be "individual differences" in formatting, but hand in 3
pages that have the info contained in this example.
Lecture 4: Quantitative Genetics: Breeder's
equation. Selection differential vs the selection
gradient: towards a more general formulation.
Frequency-dependent selection and game theory. Overheads for Lecture6
Excel Population/Quantitative-Genetic exercise #2 due 24
Sept 2013. There will be "individual differences" in
formatting, but hand in 3 pages (or more) that have the info
contained in this example.
For today's meeting, read: Maynard Smith's chapter on the basic
game-theory model (highly recommended)
Or go here for the whole
book, which is available online to IU students.
Suggested reading Kirkpatrick
(1982) in Evolution
on the evolution of female choice. Suggested
(1982) in Nature. Female choice in
Lecture 16-17. Sexual selection, chase-away selection,
Overheads for Lecture
Suggested reading: Wilkinson
& Reillo 1994. Correlated selection for female
preference in stalk-eyed flies. (see also Wilkinson et al 1998,
Suggested reading. Wilkinson
et al. 1998. Nature: meiotic drive suppression in
stalk-eyed flies (see also Wilkinson & Reillo 1994)
From Fabio: More on MHC-mediated mate choice
And in salmon:
of plasticity (not covered) <------------------
overhead slides updated to here for 2013
not covered in 2013
Exercise 4, due Nov. 2nd: create an excel worksheet that
calculates epistasis for fitness, the generation of
linkage disequilibrium, and the effect of recombination
on linkage disequilibrium. Turn in three sheets,
each one for different parameter values. Use
parameters different from the ones in my examples.
is an example. Here
is another example.
And possible chalk talk on the evolution of