Ecology (L567) 2015
Department of Biology,
evolution (L567).--This is a graduate course in
evolution, focusing on the theoretical underpinnings.
Beginning in 2010, I have put more focus on Evolutionary Ecology
and Behavioral Ecology. The goal will be to complement our
graduate courses in Evolutionary Genetics. Topics will
include life-history theory, evolution of sex, sexual selection,
evolution of social behavior, evolution of virulence, the
evolutionary ecology of speciation, and macroevolution. I also
plan to cover some of the basic theory for Population Genetics and
Quantitative Genetics. Graduate standing or permission of
the instructor is required. Advanced undergraduates in Biology are
The course will be offered
again during the Fall semester of 2015. Typically, the
class meetings are part lecture and part discussion in an
Population/Quantitative-Genetic exercise #2 due 15 Sept
2015. There will be "individual differences" in
formatting, but hand in 3 pages (or more) that have the info
contained in this example.
reading for lecture 11: Salmon paper by Einum&Fleming
(2000) Nature 405:565-567.
Lecture 11: Life-history theory: size-number compromises and
geometric mean fitness. Plus discussion of Einum and
Optional reading: The
Red Queen from Bell (1982) from, The Masterpiece of
Nature: the Evolution and Genetics of Sexuality
Lecture14: Testing the ecological hypotheses (Burt and Bell),
Difficulties on theory, Pluralism. (Most of this was an
informal talk after the first exam.)
Overheads contained in "overheads for lecture 13-14" above.
Optional reading: Pluralistic hypotheses The
ratchet and the red queen in Nature 1994 Lecture
14-15. Epistasis, Linkage Disequiblium,
Recombination, Runaway sexual selection
Suggested reading: Wilkinson
& Reillo 1994. Correlated selection for female
preference in stalk-eyed flies. (see also Wilkinson et al 1998,
Suggested reading. Wilkinson
et al. 1998. Nature: meiotic drive suppression in
stalk-eyed flies (see also Wilkinson & Reillo 1994)
From Fabio: More on MHC-mediated mate choice
And in salmon:
Optional reading: the Price
Equation by A. Gardner
Optional: for a derivation of the Price equation see (Frank, S.
2012) J. Evol. Biol. 25:1002-1019
Optional: on the relationship between group selection and kin
et al PNAS
Lecture23: The ecology of speciation I Overheads for lecture23
Exercise 4, due Nov. 2nd: create an excel worksheet that
calculates epistasis for fitness, the generation of
linkage disequilibrium, and the effect of recombination
on linkage disequilibrium. Turn in three sheets,
each one for different parameter values. Use
parameters different from the ones in my examples.
is an example. Here
is another example.