Mailing Address:
Department of Biology
142 Jordan Hall
Indiana University
1001 East Thrid Street
Bloomington, IN 47405 USA
FAX: 812.855.6705
Office: 812.855.7443
e-mail: vnolan@indiana.edu
Center for the Integrative Study of Animal Behavior (CISAB)
Val Nolan Jr.
Professor Emeritus of Biology and of Law
Professor Nolan studies the ecology and behavior of birds, principally passeriforms. His current interests are mating systems and parental behavior and the hormonal regulation of these, natal dispersal and the development of site attachment in both migratory and nonmigratory passerines, dominance, and differential migration and the differential winter distribution that such migration produces. He works almost entirely with free-living birds.
Representative Publications:
Enstrom, D.A., Soenksen, M., Ziegenfus, C., Nolan, V. Jr., and E.D. Ketterson. 200x. Natural variation in testosterone affects male attractiveness in the dark-eyed junco. Animal Behaviour, submitted, under revision.
Reed, W.L, Clark, M.E., Parker, P.G., Raouf, S.A., Arguedas, N., Monk, D.S, Snajdr, E., Nolan, V. Jr., and E.D. Ketterson. 2006. Physiological effects on demography: A long-term experimental study of testosterone's effects on fitness. American Naturalist, tentatively accepted.
Ketterson, E.D., Nolan, V. Jr., and M. Sandell. 2005. Testosterone in females: mediator of adaptive traits, constraint on the evolution of sexual dimorphism, or both? American Naturalist 166: S85-S98.
McGlothlin, J. W., P. G. Parker, Nolan, V. Jr., and E. D. Ketterson. 2005. Correlational selection leads to genetic integration of body size and an attractive plumage trait in dark-eyed juncos. Evolution 59:658-671.
McGlothlin, J., Neudorf, D.L., Casto, J.M., Nolan, V. Jr., and E.D. Ketterson. 2004. Elevated testosterone reduces choosiness in female dark-eyed juncos (Junco hyemalis): evidence for a hormonal constraint on sexual selection? Proceedings Royal Society, London B 271:1377-1384.
Clotfelter, E.D., O’Neal, D.M., Gaudioso, J.M, Casto, JM, Parker-Renga, I.M., Snajdr, E.A., Duffy, D.L., Nolan, V. Jr., and E.D. Ketterson. 2004. Consequences of elevating plasma testosterone in females of a socially monogamous songbird: evidence of constraints on male evolution? Hormones and Behavior 46:171-178.
Wolf, W.L., Casto, J.M., Nolan, V. Jr., and E. D. Ketterson. 2004. Female ornamentation and male mate choice in dark-eyed juncos. Animal Behaviour 67 (1): 93-102.
Clotfelter, E.D., Schubert, K.A., Nolan, V. Jr., and E.D. Ketterson 2003. Mouth colour signals thermal stress in nestling dark-eyed juncos (Junco hyemalis). Ethology 109:171-182.
Neudorf, D.L., Ziolkowski, D.J., Jr, Nolan V. Jr., and E.D. Ketterson. 2002. Testosterone manipluation of male attractiveness has no detectable effect on female home-range size and behavior during the fertile period. Ethology 108:713-726.
Nolan, V. Jr., Ketterson, E.D., Cristol, D.A., Rogers, C.M., Clotfelter, E.D., Titus, R., Schoech, S.J., and E. Snajdr. . 2002. Dark-eyed Junco (Junco hyemalis). In The Birds of North America, No. 716. (A. Poole and F. Gill, eds), 44p. The Birds of North America, Inc., Philadelphia, PA.
Casto, J. M., Nolan, V. Jr., and E. D. Ketterson. 2001. Steroid hormones and immune function: experimental studies in wild and captive dark-eyed juncos (Junco hyemalis). American Naturalist 157: 408-420.
Clotfelter, E. D., Nolan, V. Jr., Ketterson, E. D. 2001. The effects of experimentally elevated testosterone and food deprivation on food consumption and prey size preferences in male dark-eyed juncos (Junco hyemalis, Emberizidae: Passeriformes). Ethology 107: 439-449.
Grindstaff, J.L., Buerkle, C.M., Casto, J.M., Nolan, V. Jr., and E.D. Ketterson. 2001. Offspring sex ratio is unrelated to male attractiveness in dark-eyed juncos (Junco hyemalis). Behavioral Ecology and Sociobiology 50:312-316.
Ketterson, E. D., Nolan, V. Jr., Casto, J. M., Buerkle, C. A., Clotfelter, E., Grindstaff, J. L., Jones, K. J., Lipar, J. L., McNabb, F. M. A., Neudorf, D. L., Parker-Renga, I., Schoech, S. J., and E. Snajdr. 2001. Testosterone, phenotype, and fitness: a research program in evolutionary behavioral endocrinology. Pp. 19-40 in A. Dawson and C. M. Chaturvedi (eds.), Avian Endocrinology. Narosa Publishing House, New Delhi, India.
Nolan, V. Jr. and C. F. Thompson (eds.). 2001. Current Ornithology Vol. 16. Plenum, New York.
Nolan, V. Jr., Ketterson, E. D., and C. F. Thompson (eds.). 1999. Current Ornithology Vol. 15. Plenum, New York.
Nolan, V., Jr., E. D. Ketterson, and C. A. Buerkle. 1999. Prairie Warbler (Dendroica discolor). In The Birds of North America, No. 455 (A. Poole and F. Gill, eds.). The Birds of North American, Inc., Philadelphia, PA.
C. Ray Chandler, Ellen D. Ketterson, and Val Nolan Jr. 1997. Effects of testosterone on use of space by male dark-eyed juncos when their mates are fertile. Animal Behaviour 54:543-549.
V. Nolan Jr., E.D. Ketterson, and C.F. Thompson (Eds.) 1997. Current Ornithology, Vol. 14 (New York: Plenum Press).
Russell C. Titus, C. Ray Chandler, Ellen D. Ketterson, and Val Nolan Jr. 1997. Song rates of dark-eyed juncos do not increase when females are fertile. Behav. Ecology & Sociobiology 41:165-169.
Torgeir S. Johnsen, James D. Hengeveld, James L. Blank, Ken Yasukawa & Val Nolan Jr. 1996. Epaulet brightness and condition in female red-winged blackbirds. The Auk 113:356-362.
V. Nolan Jr. and E.D. Ketterson (Eds.). 1996. Current Ornithology Vol. 13 (New York: Plenum Press).
E.D. Ketterson and V. Nolan Jr. 1994. Male parental behavior in birds. Annual Review of Ecology and Systematics 25:601-628.
Val Nolan Jr., Ellen D. Ketterson, Charles Ziegenfus, Daniel P. Cullen, and C. Ray Chandler. 1992. Testosterone and avian life histories: Effects of experimentally elevated testosterone on prebasic molt and survival in male dark-eyed juncos. The Condor 94:364-370.
Val Nolan Jr. and Ellen D. Ketterson. 1990. Timing of autumn migration and its relation to winter distribution in dark-eyed juncos. Ecology 71:1267-1278.
Blank, J. and V. Nolan Jr. 1983. Offspring sex ratio in red-winged blackbirds is determined by maternal age. Publications of National Academy of Science 80:6141-6145.
Carey, M. and V. Nolan Jr. 1979. Population dynamics of the indigo bunting and the evolution of avian polygyny. Evolution 33:1180-1192.
Val Nolan Jr. 1978. The behavior and ecology of the prairie warbler Dendroica discolor. Ornithological Monographs 26:1-595.
Carey, M. and V. Nolan Jr. 1975. Polygyny in indigo buntings: a hypothesis tested. Science 190:1296-1297.